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| The
Structure of a single barley floret and bracts (spikelet). |
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The floral structure
of the barley plant is the spike, or, as it is sometimes referred
to, the head or ear (6). The spike rests at the tip of each stem,
also known as a tiller. Several quality variations between six-and
two-row barley are directly attributable to spike morphology, so
an understanding of the differences is meaningful.
The central
axis of the spike is known as the rachis. The rachis is composed
of nodes and internodes, with spikelets that can later develop into
kernels attached at the rachis nodes. A barley spikelet comprises
an individual floret with surrounding bracts. The barley floret
is "perfect," meaning that it contains both male (stamen) and female
(pistil) floral components; barley is self-pollinating. Following
pollination, the fertilized ovary develops into the embryo (germ)
and endosperm of the kernel, while the bracts form the husk of the
mature kernel.
In
both two- and six-row barley, each individual node of the rachis
has three spikelets, but the fertility (or sterility) of the florets
differs in each type. In six-row barley, all three spikelets (per
rachis node) contain a fertile floret. These florets develop into
kernels and thus each rachis node in the mature spike of six-row
barley has three kernels. When the rachis is viewed from one side,
there appears to be three rows of kernels. Kernels occur at nodes
on both sides of the zigzag rachis, however, so it looks like six
rows. This is perhaps most easily seen when viewed downward from
the top of the spike. In two-row barley, only the central floret
is fertile and will develop into a kernel; the lateral spikelets
are sterile, leaving alternating single kernels on opposite sides
of the rachis, for a total of two rows. Barley has multiple stems
(tillers) per plant, with many of the stems producing spikes. Two-row
barley plants generally have more spikes per plant, but the number
of tillers is greatly influenced by environmental conditions.
An
additional characteristic used to distinguish malting barley types
is the color of the aleurone (3). The aleurone layer is directly
under the husk tissues and largely surrounds the endosperm. It is
of extreme importance in malting because it is a major site of enzyme
synthesis (7). When certain phenolic pigments are present in the
aleurone, they may give the dehusked (pearled) grain a black, violet,
purple, blue, or green appearance. It should be noted that these
aleurone pigments do not have an easily identified contribution
to beer color, but are thought to impact flavor. But again, this
impact is not easily quantified. Absence of pigment results
in a white aleurone. Blue aleurone color, which is controlled by
a single gene, was once quite common in North American six-row cultivars.
In the Canadian grain trade, the blue aleurone trait was once used
as a marker to distinguish six-row malting cultivars (blue) from
six-row feed cultivars. For the most part, blue aleurone barleys
have fallen out of favor with North American brewers, and little
blue aleurone is currently produced. All North American two-row
barley cultivars have white aleurone.
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